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I think we are seeing a blastodisc maybe. My bullseye were more dramatic than this.

Thread 'Are these eggs fertile?' https://www.backyardchickens.com/threads/are-these-eggs-fertile.1521898/ I may agree on this assessment. Follow the link above, I think your eggs look very similar. It's always hard to tell from pictures but I don't see a defined bullseye.

I can't believe the storage temperature would greatly affect your outcome. I have successfully incubated eggs I pulled it out of the refrigerator that were over a week old. I suspect a fertility issue. We can look at images of what looks like a bullseye, but that doesn't necessarily mean that...